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note
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We thank P. Vitousek, R. Naylor, R. Kelly, and G. Daily for extended discussions on these topics, G. P. Robertson and two anonymous reviewers for thoughtful reviews of earlier drafts, and K. McElwain and B. Techau for assistance with manuscript preparation. Partially supported by grants to W.J.P. (NASA-NAGW-2662, NSF-BSR-9011659, NIH-1-R01-HD33554, and EPA-R824993-01-0), P.A.M. (NASA-NAGW-3869 and USDA-94-37101-0437), and A.G.P. (NSF-9407919 and USDA-95-33120-1877).
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