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2 (equal to 95.2% exclusion efficiency). Lateral movement traps, located outside and inside of the lateral movement fence, indicated a similar efficiency (95.0%). Primary production in the headwater streams at Coweeta is very low (4).
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Examples of taxa that feed on material buried in the substratum are sericostomatid caddisfly larvae, Fattigia [R. Wagner, Hydrobiologia 224, 65 (1991)]; Diptera, Molophilus (16); and Oligochaeta. We extracted gut contents of larvae and measured projected areas of various food resources by using a compound microscope, camera lucida, and digitizing tablet [ J. B. Wallace et al., Arch. Hydrobiol. Suppl. 74, 423 (1987)]. Guts of some specimens contained >60% woody tissue. Others have reported woody tissue as common in guts of insects, such as Lipsothrix [ T. L. Dudley and N. H. Anderson, Freshwater Biol. 17, 437 (1987)] and Paraleptophlebia, Tipula, and Sciaridae [ C. R. D. Pereira et al., Melanderia 39, 23 (1982)].
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Examples of taxa that feed on material buried in the substratum are sericostomatid caddisfly larvae, Fattigia [R. Wagner, Hydrobiologia 224, 65 (1991)]; Diptera, Molophilus (16); and Oligochaeta. We extracted gut contents of larvae and measured projected areas of various food resources by using a compound microscope, camera lucida, and digitizing tablet [ J. B. Wallace et al., Arch. Hydrobiol. Suppl. 74, 423 (1987)]. Guts of some specimens contained >60% woody tissue. Others have reported woody tissue as common in guts of insects, such as Lipsothrix [ T. L. Dudley and N. H. Anderson, Freshwater Biol. 17, 437 (1987)] and Paraleptophlebia, Tipula, and Sciaridae [ C. R. D. Pereira et al., Melanderia 39, 23 (1982)].
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Examples of taxa that feed on material buried in the substratum are sericostomatid caddisfly larvae, Fattigia [R. Wagner, Hydrobiologia 224, 65 (1991)]; Diptera, Molophilus (16); and Oligochaeta. We extracted gut contents of larvae and measured projected areas of various food resources by using a compound microscope, camera lucida, and digitizing tablet [ J. B. Wallace et al., Arch. Hydrobiol. Suppl. 74, 423 (1987)]. Guts of some specimens contained >60% woody tissue. Others have reported woody tissue as common in guts of insects, such as Lipsothrix [ T. L. Dudley and N. H. Anderson, Freshwater Biol. 17, 437 (1987)] and Paraleptophlebia, Tipula, and Sciaridae [ C. R. D. Pereira et al., Melanderia 39, 23 (1982)].
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2 during the third year of exclusion) constituted a large residual source of terrestrially derived organic matter available to invertebrates throughout the study. Furthermore, inputs of groundwater dissolved organic carbon were not curtailed.
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2 during 3 years of exclusion. Export of fine particles has been measured continuously in both streams since early 1985 [T. F. Cuffney and J. B. Wallace, Can. J. Fish. Aquat. Sci. 45, 2010 (1988)]. Fine organic particle export in the exclusion stream did not decrease relative to the reference stream until the third year of exclusion ( J. B.Wallace et al., unpublished data).
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2 during 3 years of exclusion. Export of fine particles has been measured continuously in both streams since early 1985 [T. F. Cuffney and J. B. Wallace, Can. J. Fish. Aquat. Sci. 45, 2010 (1988)]. Fine organic particle export in the exclusion stream did not decrease relative to the reference stream until the third year of exclusion ( J. B.Wallace et al., unpublished data).
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We thank E. F. Benfield, A. C. Benke, A. D. Huryn, M. D. Hunter, J. Hutchens, A. Rosemond, and E. Rosi for constructive comments and W. T. Swank, B. Keeley, and other personnel from the Coweeta Hydrologic Laboratory for on-site support. This research was supported by NSF.
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