Pterosaur tracks and the terrestrial ability of pterosaurs

Lethaia

Volumn 29, Issue 4, 1996, Pages 373-386

  Unwin, David M ^a  

^a UNIVERSITY OF BRISTOL   (United Kingdom)

Author keywords

Functional morphology; Jurassic; Palaeoichnology; PTERAICHNUS; Pterosaur; Vertebrate

Indexed keywords

ANIMALIA; CROCODYLIDAE (ALL CROCODILES); PTERAICHNUS; VERTEBRATA;

FUNCTIONAL MORPHOLOGY; JURASSIC; PTEROSAUR; TRACKWAY;

EID: 0030405383     PISSN: 00241164     EISSN: None     Source Type: Journal    
DOI: 10.1111/j.1502-3931.1996.tb01673.x     Document Type: Article

References (87)

1. Abel, O. 1925: On a skeleton of Pterodactylus antiquus from the lithographic shales of Bavaria, with remains of skin and musculature. American Museum Novitates 192, 1-12.
2. Anonymous, 1973: Track of the pterosaur: Probable oldest evidence. Science News 104, 85.
3. Baird, D. 1954: Chirotherium lulli, a pseudosuchian reptile from New Jersey. Bulletin of the Museum of Comparative Zoology (Harvard University) 11, 165-192.
4. Bakhurina, N.N. 1986: Letayushchie yashcheny. [Flying reptiles.] Priroda (1986) 7, 27-36. [In Russian].
5. Bakhurina, N.N. 1992: Pterosaurs from continental deposits of Middle Asia, Mongolia and China. Abstracts of the Palaeontological Association Annual Conference, Southampton, Palaeontology Newsletter 16, 11.
6. Bakhurina, N.N. 1993: Early Cretaceous pterosaurs from western Mongolia and the evolutionary history of the Dsungaripteroidea. Journal of Vertebrate Paleontology 13:3, Supplement, 24A.
7. Bakhurina, N.N. & Unwin D.M. 1995: A survey of pterosaurs from the Jurassic and Cretaceous of the former Soviet Union and Mongolia. Historical Biology 10, 197-245.
8. Bennett, S.C. 1990: A pterodactyloid pterosaur from the Santana Formation of Brazil: implications for terrestrial locomotion. Journal of Vertebrate Paleontology 10, 80-85.
9. Bennett, S.C. 1992: Reinterpretation of problematical tracks at Clayton Lake State Park, New Mexico: not one pterosaur, but several crocodiles. Ichnos 2, 37-42.
10. Bramwell, C.D. & Whitfield, G.R. 1974: Biomechanics of Pteranodon. Philosophical Transactions of the Royal Society of London (B) 267, 503-581.
11. Broili, F. 1927: Ein Exemplar von Rhamphorhynchus mit Resten von Schwimmhaut. Sitzungs-Berichte der Bayerischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Abteilung, 29-48.
12. Broili, F. 1936: Weitere Beobachtungen an Ctenochasma. SitzungsBerichte der Bayerischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Abteilung, 137-156.
13. Broili, F. 1938: Beobachtungen an Pterodactylus. Sitzungs-Berichte der Bayerischen Akademie der Wissenschaften, mathematisch-naturwissen-schaftliche Abteilung, 139-154.
14. Cadèe, G.C. 1990: Feeding traces and bioturbation by birds on a tidal flat, Dutch Wadden Sea. Ichnos 1, 23-30.
15. Caster, K.E. 1941: Trails of Limulus and supposed vertebrates from the Solnhofen lithographic limestone. Pan American Geologist 76, 241-258.
16. Caster, K.E. 1957: Problematica. Memoirs of the Geological Society of America 67, 1025-1032.
17. Conrad, K., Lockley, M.G. & Prince, N.K. 1987: Triassic and Jurassic vertebrate-dominated trace fossil assemblages of the Cimarron Valley region: Implications for paleoecology and biostratigraphy. New Mexico Geological Society Guidebook, 38th Field Conference, 127-138.
18. Currie P.J. 1989: Dinosaur footprints of western Canada. In Gillette, D.D. & Lockley, M.G. (eds): Dinosaur Tracks and Traces, 293-300. Cambridge University Press, Cambridge.
19. Delair, J. B. 1963: Notes on Purbeck fossil footprints, with descriptions of two hitherto unknown forms from Dorset. Proceedings of the Dorset Natural History and Archaeological Society 84, 92-100.
20. Döderlein, L. 1929: Ein Pterodactylus mit Kehlsack und Schwimmhaut. Sitzungs-Berichte der Bayerischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse 65-76.
21. Ensom, P. 1984: Purbeckopus pentadactylus Delair. Proceedings of the Dorset Natural History and Archaeological Society 105, 166.
22. Figuier, L. 1863: La Terre avant le deluge. 448 pp. Paris.
23. Fregenal-Martinez, M.A. & Moratalla, J.J. 1995: Paleoichnology. In Nieves Meléndez, M. (ed.): Las Hoyas. A lacustrine Konservat-Lager-stätte, Cuenca, Spain, 71-75. Field Trip Guide Book, II International Symposium on Lithographic Limestones, Cuenca, Spain 1995.
24. Gillette, D.D. & Thomas, D.A. 1989: Problematical tracks and traces of Late Albian (Early Cretaceous) Age, Clayton Lake State Park, New Mexico, USA. In Gillette, D.D. & Lockley, M.G. (eds): Dinosaur Tracks and Traces, 337-342. Cambridge University Press, Cambridge.
25. Hazlehurst, G.A. & Rayner, J.M.V. 1992: Flight characteristics of Triassic and Iurassic Pterosauria: an appraisal based on wing shape. Paleobiology 18, 447-463.
26. Hunt, A.P., Lockley, M.G., Hups, K. & Schultz, R. 1995: Jurassic vertebrate paleontology of Cactus Park, west-central Colorado. Geological Society of America, Abstracts with Programs, Rocky Mountains Section, 27:4, 15.
27. Längsten, W. Jr. 1974: Non-mammalian Comanchean tetrapods. Geoscience and Man 8, 77-102.
28. Leonardi, G. 1987: Glossary and manual of Tetrapod footprint palaeoichnology. Departamento Nacional de Producao. 75 pp.
29. Lockley, M.G. 1991: Tracking Dinosaurs. 238 pp. Cambridge University Press, Cambridge.
30. Lockley, M.G., Yang, S-Y., Matsukawa, M., Fleming, F. & Lim, S.-K. 1992: The track record of Mesozoic birds: evidence and implications. Philosophical Transactions of the Royal Society of London 336, 113-134.
31. Lockley, M.G. & Hunt, A.P. 1995: Dinosaur Tracks and Other Fossil Footprints of the Western United States. 338 pp. Columbia University Press, New York, N.Y.
32. Lockley, M.G., Logue, T.J., Moratalla, J.J., Hunt, A.P., Schultz, R.J. & Robinson, J.W. 1995: The fossil trackway Pteraichnus is pterosaurian, not crocodilian: implications for the global distribution of pterosaur tracks. Ichnos 4, 7-20.
33. Logue, T.J. 1977: Preliminary investigations of pterodactyl tracks at Al-cova, Wyoming. Wyoming Geological Association. Earth Science Bulletin 10, 29-30.
34. Logue, T.J. 1994: Alcova, Wyoming tracks of Pteraichnus saltwashensis made by pterosaurs. Geological Society of America, Abstracts with Program, South Central Region 26, 10.
35. Mazin, J.-M., Hantzpergue, P., Lafaurie, G. & Vignaud, P. 1995: Des pistes de ptérosaures dans le Tithonien de Crayssac (Quercy, France). Comptes rendus de l'Académie des Sciences de Paris 321, 417-424.
36. Meyer, H. 1859: Zur Fauna der Vorwelt. Vierte Abt.: Reptilien aus dem lithographischen Schiefer des Jura in Deutschland und Frankreich, 1-84. Frankfurt-am-Main.
37. [Moratalla, J.J. 1993: Restos indirectos de dinosaurios del registro español: Paleoicnologia de la Cuenca de Cameros Jurassico superior - Cretacico inferior y Paleoologia del Cretacico superior. PhD thesis. Universidad Autonoma de Madrid.]
38. Oppel, A. 1862: Über Fahrten in lithographischen Schiefer. Mitteilungen Museum Bayerische Staatssammlung 1, 121-125.
39. Owen, R. 1870: Monograph on the Fossil Reptilia of the Liassic Formations. I. Part III, 41-81. The Palaeontographical Society, London.
40. Padian, K. 1983a: A functional analysis of flying and walking in pterosaurs. Paleobiology 9, 218-239.
41. Padian, K. 1983b: Osteology and functional morphology of Dimorphodon macronyx (Buckland) (Pterosauria: Rhamphorhynchoidea) based on new material in the Yale Peabody Museum. Postula 189, 1-44.
42. Padian, K. 1984: The origins of pterosaurs. In Reif, W.-E. & Westphal, F. (eds): Third Symposium on Mesozoic Terrestrial Ecosystems Short Papers, 163-168. Attempto, Tübingen.
43. Padian, K. 1985: The origins and aerodynamics of flight in extinct vertebrates. Palaeontology 28, 413-433.
44. Padian, K. 1987: The case of the bat-winged pterosaur. Typological taxonomy and the influence of pictorial representation on scientific perception. In Czerkas, S.J. & Olson, E.C. (eds): Dinosaurs Past and Present 2, 65-81. Natural History Museum of Los Angeles County and University of Washington Press, Seattle, Wash.
45. Padian, K. 1988: The flight of pterosaurs. Natural History 97:12, 58-65.
46. Padian, K. 1991: Pterosaurs: were they functional birds or functional bats? in Rayner, J.M.V. & Wootton, R. J. (eds): Biomechanics in evolution, 146-160. Seminar Series of the Society for Experimental Biology 36. Cambridge University Press, Cambridge.
47. Padian, K. & Olsen, P.E. 1984: The fossil trackway Pteraichnus: not pterosaurian, but crocodilian. Journal of Paleontology 58, 178-184.
48. Padian, K. & Rayner, J.M.V. 1993: The wings of pterosaurs. American Journal of Science 293A, 91-166.
49. Padian, K., Ricqlès, A.J. de, & Horner, J.R. 1995: Bone histology determines identification of a new fossil taxon of pterosaur (Reptilia: Archosauria). Comptes rendus de l'Académie des Sciences de Paris 320, 77-84.
50. Parker, L. & Balsley, J. 1989: Coal mines as localities for studying trace fossils. In Gillette, D.D. & Lockley, M.G. (eds): Dinosaur Tracks and Traces, 353-359. Cambridge University Press, Cambridge.
51. Paul, G.S. 1987: Pterodactyl habits - real and radio controlled. Nature 328, 481.
52. Pennycuick, C.J. 1986: Mechanical constraints on the evolution of flight. In Padian, K. (ed.): The Origin of Birds and the Evolution of Flight, 83-98. Memoirs of the California Academy of Sciences 8.
53. Prince, N.K. & Lockley, M.G. 1989: The sedimentology of the Purgatoire Tracksite Region, Morrison Formation, of South-eastern Colorado. In Gillette, D.D. & Lockley, M.G. (eds): Dinosaur Tracks and Traces, 155-163. Cambridge University Press, Cambridge.
54. Reineck, H. & Howard, I.D. 1978: Alligatorfahrten. Natur und Museum 108, 10-15.
55. Sharov, A.G. 1971: Novye letayushchie reptilii iz mezozoya Kazakhstana i Kirgizii. [New flying reptiles from the Mesozoic of Kazakhstan and Kirghizia.] Trudy Paleontologicheskogo lnstituta AN SSSR 130, 104-113. (In Russian.)
56. Stieler, C. 1922: Neuer Rekonstruktionsversuch eines liassischen Flugsauriers. Naturwissenschaften Wochenschrift NF 21:20, 273-280.
57. Stokes, W.L. 1957: Pterodactyl tracks from the Morrison Formation. Journal of Paleontology 31, 952-954.
58. Stokes, W.L. 1978: Animal tracks in the Navajo-Nugget Sandstone. Contributions to Geology of the University of Wyoming 16, 103-107.
59. Stokes, W.L. & Madsen, I.H. Jr. 1979: Environmental significance of pterosaur tracks in the Navajo Sandstone (Jurassic) Grand County, Utah. Brigham Young University Geological Studies 26, 21-26.
60. Stricklin, F.L. Jr. & Amsbury, D.L. 1974: Depositional environment on a low-relief carbonate shelf, middle Glen Rose Limestone, central Texas. Geoscience and Man 8, 53-66.
61. Thulborn, T. 1990: Dinosaur Tracks. 410 pp. Chapman & Hall, London.
62. Unwin, D.M. 1986: Tracking the dinosaurs. Geology Today 2, 168-169.
63. Unwin, D.M. 1987a: Pterosaur locomotion. Joggers or waddlers? Nature 327, 13-14.
64. Unwin, D.M. 1987b: Reconstructing extinct animals: did pterosaurs walk or waddle? Anima 180, 39-43. (In Japanese.)
65. Unwin, D.M. 1988: New remains of the pterosaur Dimorphodon (Pterosauria: Rhamphorhynchoidea) and the terrestrial ability of early pterosaurs. Modern Geology 13, 57-68.
66. Unwin, D.M. 1989: A predictive method for the identification of vertebrate ichnites and its application to pterosaur tracks. In Gillette, D.D. & Lockley, M.G. (eds): Dinosaur Tracks and Traces, 259-274. Cambridge University Press, Cambridge.
67. Unwin, D.M. & Bakhurina, N.N. 1994: Sordes pilosus and the nature of the pterosaur flight apparatus. Nature 371, 62-64.
68. Valenzuela, M., Garcia-Ramos, J.-C. & Suarezn de Centi, C. 1988: Las huellas de dinosaurios de entorno de Ribadesella. Central Lechera Asturiana. 35 pp.
69. Wellnhofer, P. 1970: Die Pterodactyloidea (Pterosauria) der Oberjura Plattenkalke Süddeutschlands. Abhandlungen der Bayerischen Akademie der Wissenschaften zu München, Mathematisch-Naturwissen-schaftliche Klasse 141, 1-133.
70. Wellnhofer, P. 1975a: Die Rhamphorhynchoidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands. I. Allgemeine Skelettmorphologie. Palaeontographica A 148, 1-33.
71. Wellnhofer, P. 1975b: Die Rhamphorhynchoidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands. HI. Palökologie und Stammesgeschichte. Palaeontographica A 149, 1-30.
72. Wellnhofer, P. 1975c: Die Rhamphorhynchoidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands. II. Systematische Beschreibung. Palaeontographica A 148, 132-186.
73. Wellnhofer, P. 1978: Handbuch der Paläoherpetologie. Teil 19, Pterosauria. 82 pp. Gustav Fischer, Stuttgart.
74. Wellnhofer, P. 1980: Flugsaurier. Neue Brehm-Bücherei 534. 135 pp. Ziemsen, Wittenburg-Lutherstadt.
75. Wellnhofer, P. 1983: Solnhofener Plattenkalk: Urvögel und Flugsaurier. 59 pp. Freunde des Museums beim Solenhofer Aktien-Verein, Maxberg.
76. Wellnhofer, P. 1985: Neue Pterosaurier aus der Santana Formation (Apt.) der Chapada do Araripe, Brasilien. Palaeontographica A 187, 105-182.
77. Wellnhofer, P. 1987: Die Flughaut von Pterodactylus (Reptilia. Pterosauria) am Beispiel des Wiener Exemplares von Pterodactylus kochi (Wagner). Annalen des Naturhistorischen Museums Wien 88A, 149-162.
78. Wellnhofer, P. 1988: Terrestrial locomotion in pterosaurs. Historical Biology 1, 3-16.
79. Wellnhofer, P. 1991a: The Illustrated Encyclopedia of Pterosaurs. 192 pp. Salamander Books, London.
80. Wellnhofer, P. 1991b: Weitere Pterosaurierfunde aus der Santana-Formation (Apt.) der Chapada do Araripe, Brasilien. Palaeontographica A 215, 43-101.
81. Wellnhofer, P. 1991c: Santana Formation pterosaurs. In Maisey, J.G. (ed.): Santana Fossils, 351-370. T.F.H. Publications Inc, Neptune City, N.J.
82. Wellnhofer, P. & Vahldiek, B.W. 1986: Ein Flugsaurier-Rest aus dem Posidonienschiefer (Unter-Toarcian) von Schandeiah bei Braunschweig. Paläontologische Zeitschrift 60:3/4, 329-340.
83. [West, E.S. 1978: Biostratigraphy and Paleoecology of the Lower Morrison Formation of Cimarron County, Oklahoma. 61 pp. Unpublished PhD Dissertation.]
84. Williston, S.W. 1902: On the skeleton of Nyctodactylus, with restoration. American Journal of Anatomy 1, 297-305.
85. Wright, J.L., Unwin, D.M., Lockley, M.G. & Rainforth, E. (in press): Pterosaur tracks from the Purbeck Formation of Dorset, England. Proceedings of the Geologist's Association.
86. Young, C.C. 1964: On a new pterosaurian from Sinkiang, China. Vertebrata Palasiatica 8, 221-255.
87. Young, C.C. 1973: [Wuerho pterosaurs.] Special Publication of the Institute of Vertebrate Palaeontology and Palaeoanthroplology, Academica Sinica, No. 11, 18-34. (In Chinese.)