Generic revision of the New World tribe Hoplophorionini (Hemiptera: Membracidae: Membracinae)

Systematic Entomology

Volumn 21, Issue 4, 1996, Pages 295-342

  Mckamey, Stuart H ^a,c   Deitz, Lewis L ^b  

^a UNIVERSITY OF CONNECTICUT   (United States)

^b NORTH CAROLINA STATE UNIVERSITY   (United States)

^c UNIVERSITY OF COPENHAGEN   (Denmark)

Author keywords

[No Author keywords available]

Indexed keywords

HEMIPTERA; MEMBRACIDAE;

TREEHOPPERS;

EID: 0030319490     PISSN: 03076970     EISSN: None     Source Type: Journal    
DOI: 10.1111/j.1365-3113.1996.tb00602.x     Document Type: Article

References (123)

1. Abud, A.J. (1976) [Report of Umbonia crassicornis in the Dominican Republic]. Universidad Autónoma de Santo Domingo, Departamento de Biología, Naturalista, 16/76. [1 p.].
2. Amyot, C.J.B. & Serville, [J.G.].A (1843) Deuxième partie. Homoptères. Homoptera Latr. Histoire Naturelle des Insectes Hémiptères, pp. 455-676. Libraire Encyclopédique de Roret, Paris, France.
3. Arnett, R.H., Sammuelson, G.A. & Nishida, G.M. (1993) The Insect and Spider Collections of the World, 2nd edn. (Flora & Fauna Handbook no. 11.) Sandhill Crane Press, Inc., Gainesville, Florida.
4. Ballou, C.H. (1935) Insect notes from Costa Rica in 1934. Insect Pest Survey Bulletin, 15, 163-212.
5. Ballou, C.H. (1936a) Insectos observados durante el año 1934. Centro Nacional de Agricultura Boletín, 20, 1-60.
6. Ballou, C.H. (1936b) Insect notes from Costa Rica in 1935. Insect Pest Survey Bulletin, 16 (Suppl. to no. 9), 437-497.
7. Beamer, R.H. (1930) Maternal instinct in a membracid (Platycotis vittata) (Homop.). Entomological News, 41, 330-331, plate 35.
8. Bodenheimer, F.S. (1951) Insects as Human Food, a Chapter of the Ecology of Man. Dr W. Junk, The Hague.
9. Bondar, G, (1939) Insetos daninhos e parasitas do caucau na Bahia. Instituto de Cacau da Bahia Boletin Tecnico, 5 [i-vi], 1-112.
10. Brach, V. (1975) A case of active brood defense in the thornbug, Umbonia crassicornis (Homoptera: Membracidae). Bulletin of the Southern California Academy of Science, 74, 163-174.
11. Brown, R.L. & Eads, C.O. (1965) A technical study of insects affecting the oak tree in southern California. California Agricultural Experiment Station Bulletin, 810, 1-105.
12. Burmeister, H.C.C. (1835) Schnabelkerfe. Rhynchota. Handbuch der Entomologie, 2(1), 1-396.
13. Butcher, F.G. (1953) Unusual abundance of the tree-hopper Umbonia crassicornis A & S [sic]. Florida Entomologist, 36, 57-59.
14. Callan, E., Mc.C. (1976) Observations on the nesting behavior and prey of gorytine wasps in Trinidad (Hymenoptera, Sphecidae). Psyche, 83, 324-335.
15. Camacho, A.A. (1944) El periquito del aguacate. Fitófilo (Boletín de la Oficina Fitosanitaria. San-Jacinto, México, D.F.), 3 (4), 3-54.
16. Carpenter, J.M. (1993) Random cladistics. Cladistics, 8, 147-153.
17. Castner, J.L. (1985) Oak treehoppers (Platycotis vittata) in Florida. BioScience, 35 (11), cover, 684.
18. Chapin, E.A. (1925) Food habits of the vireos; a family of insectivorous birds. United Stales Department of Agriculture Bulletin, 1355, 1-42.
19. Chellman, C.W. (1978) Pests and Problems of South Florida Trees and Palms. Florida Department of Agriculture & Consumer Services, Divison of Forestry [Gainesville].
20. Christensen, J.R. (1943) Lista de Membracidae encontrados en la Argentina y algunos paises limitrofes. Revista de la Sociedad Entomológica de Argentina, 11 (5), 440-445.
21. de Conconi, J.R.-E. & Bourges-Rodriguez, H. (1977) Valor nutritivo de ciertos insectos comestibles de México y lista de algunos insectos comestibles del mundo. Annales del Instiluto de Biología de la Universidad Nacional Autónoma de México, 48 (ser. Zool.) (1), 165-186.
22. Cook, P.R (1955) Notes on nomenclature and variation in Platycotis (Homoptera: Membracidae). Pan-Pacific Entomologist, 31, 151-154.
23. Costa Lima, A.M., da. (1936) Homoptera. Terceiro Catalogo dos Insectos que Vivem nas Plantas do Brasil, pp. 131-136. Ministerio da Agricultura, Departamento Nacional da Producção Vegetal Escola Nacional de Agronomia, Rio de Janeiro.
24. Daniel, H. (1946) Aspectos de la lucha biológica. Universidad de Antioquía Medellin, Colombia, 78-79, 243-249.
25. Deitz, L.L. (1975) Classification of the higher categories of the New World treehoppers (Homoptera: Membracidae). North Carolina Agricultural Experiment Station Technical Bulletin, 225, [i-iv], 1-177.
26. Deitz, L.L. (1989) Bibliography of the Membracoidea (Homoptera: Aetalionidae, Biturritiidae, Membracidae, and Nicomiidae) 1956-1980. North Carolina Agricultural Research Service Technical Bulletin, 290, 1-31.
27. Deitz, L.L. & Dietrich, C.H. (1993) Superfamily Membracoidea (Homoptera: Auchenorrhyncha): I. Introduction and revised classification with new family group taxa. Systematic Entomology, 18, 287-296.
28. Deitz, L.L. & Kopp, D.D. (1987) Bibliography of the Membracoidea (Homoptera: Aetalionidae, Biturritiidae, Membracidae, and Nicomiidae) 1981-1987. North Carolina Agricultural Research Service Technical Bulletin, 284, 1-39.
29. Dietrich, C.H. (1989) Surface sculpturing of the abdominal integument of Membracidae and other Auchenorrhyncha (Homoptera). Proceedings of the Entomological Society of Washington, 91, 143-152.
30. Dietrich, C.H. & Deitz, L.L. (1991) Revision of the Neotropical treehopper tribe Aconophorini (Homoptera: Membracidae). North Carolina Agricultural Research Service Technical Bulletin, 293, 1-134.
31. Dietrich, C.H. & Deitz, L.L. (1993) Superfamily Membracoidea (Homoptera: Auchenorrhyncha): II. Cladistic analysis and conclusions. Systematic Entomology, 18, 297-311.
32. Dietrich, C.H. & McKamey, S.H. (1995) Two new Neotropical treehopper genera and investigation of the phylogeny of the subfamily Membracinae (Homoptera: Membracidae). Proceedings of the Entomological Society of Washington, 97, 1-16.
33. Dowell, R.V. & Johnson, M. (1986) Polistes major (Hymenoptera: Vespidae) predation of the treehopper, Umbonia crassicornis (Homoptera: Membracidae). Pan-Pacific Entomologist, 62, 150-152.
34. Dozier, H.L. (1920) Notes on the genus Platycotis Stål. Ohio Journal of Science, 20, 209-212, plate I.
35. Dozier, H.L. (1931) New and interesting West Indian Homoptera. American Museum Novitates, 510, 1-24.
36. Dozier, H.L. (1932) Descriptions of new trichogrammatid (Hymenoptera) egg parasites from the West Indies. Proceedings of the Entomological Society of Washington, 34, 29-37.
37. Ekkens, D. [B.] (1972) Peruvian treehopper behavior (Homoptera: Membracidae). Entomological News, 83, 257-271.
38. Essig, E.O. (1926) Insects of Western North America. Macmillian Co., New York.
39. Essig, E.O. (1958) Insects and Mites of Western North America, revised edn. Macmillian Co., New York.
40. Evans, H.E. (1966) The Comparative Ethology and Evolution of the Sand Wasps. Harvard University Press, Cambridge, Massachusetts.
41. Farris, J.S. (1969) A successive approach to character weighting. Systematic Zoology, 18, 374-385.
42. Farris, J.S. (1988) HENNIG86, version 1.5. Computer software and documentation, not paginated, distributed by author.
43. Fonseca, J.P. da. & von Diringshofen, R. (1974) Contribuição ao conhecimento dos membracídeos neotrópicos (Homoptera: Membracidae), VII. Arquivos de Instituto Biológico (São Paulo), 41, 151-160.
44. Fowler, W.W. (1894) Some new species of Membracidae. Transactions of the Royal Entomological Society of London, 1894, 415-424.
45. Funkhouser, W.D. (1951) Homoptera fam. Membracidae. Genera Insectorum, 208, 1-383.
46. Gaud, S.M. & Martorell, L.F. (1973) New insect record for Puerto Rico. Journal of Agriculture of the University of Puerto Rico, 57, 247-254.
47. Germar, E.F. (1835) Species Membracidum Musae E. F. Germari. Revue d'Entomologie, Gustave Silbermann, 3, 223-262.
48. Goding, F.W. (1926) Classification of the Membracidae of America. Journal of the New York Entomological Society, 34, 295-317.
49. Goding, F.W. (1929) The Membracidae of South America and the Antilles. IV. Subfamilies Hoplophorioninae, Darninae, Smiliinae, Tragopinae (Homoptera). Transactions of the American Entomological Society, 55, 197-330.
50. Hansen, H.J. (1900) The morphology and classification of the auchenorrhynchous Homoptera. Entomologist, 33, 334-337.
51. Haviland, M.D. (1925) The Membracidae of Kartabo, Bartica District, British Guiana, with descriptions of new species and bionomical notes. Zoologica (New York), 6, 229-290.
52. Hayward, K.J. (1942) Primera lista de insectos tucumanos perjudiciales. Estación Experimental Agrícola de Tucumán, Publicación Miscelanea, 1, 1-110.
53. Hayward, K.J. (1944) Primera lista de insectos tucumanos perjudiciales. Primer suplemento. Estación Experimental Agrícola de Tucumán, Publicación Miscelanea, 4, 1-32.
54. Howard, F.W. (1980) Attractiveness of date and coconut palms to Myndus crudus and other homopterans. Proceedings of the Florida State Horticultural Society, 93, 199-201.
55. Huelsenbeck, J.P. (1991) Tree-length distribution skewness: an indicator of phylogenetic information. Systematic Zoology, 40, 257-270.
56. Hunt, R.E. (1994) Vibrational signals associated with mating behavior in the treehopper, Enchenopa binotata Say (Hemiptera: Homoptera: Membracidae). Journal of the New York Entomological Society, 102 (2), 266-270.
57. Keese, M.C. & Wood, T.K. (1991) Host-plant mediated geographic variation in the life history of Platycotis vittata (Homoptera: Membracidae). Ecological Entomology, 16, 63-72.
58. Kirkaldy, G.W. (1901) Miscellanea Rhynchotalia. Entomologist, 34, 5-6.
59. Kirkaldy, G.W. (1904) Bibliographical and nomenclatorial notes on the Hemiptera. - No. 3. Entomologist, 37, 279-283.
60. Kopp, D.D. & Yonke, T.R. (1973) The treehoppers of Missouri: Part 1. Subfamilies Centrotinae, Hoplophorioninae, and Membracinae (Homoptera: Membracidae). Journal of the Kansas Entomological Society, 46, 42-64.
61. Laroca, S. & Sakakibara, A.M. (1976) Mutualismo entre Trigona hyalinata branneri (Apidae) e Aconophora flavipes (Membracidae). Revista Brasileira de Entomologia, 20, 71-72.
62. Loye, J.E. (1992) Ecological diversity and host plant relationships of treehoppers in a lowland tropical rainforest (Homoptera: Membracidae and Nicomiidae). Insects of Panama and Mesoamerica: Selected Studies (ed. by D. Quintero & A. Aiello), pp. 280-289. Oxford University Press, New York.
63. Mabberly, D.J. (1989) The Plant-Book: a Portable Dictionary of the Higher Plants, corrected edn. Cambridge University Press, Cambridge, England.
64. McAtee, W.L. (1918) Bird enemies of tree hoppers (Membracidae). Auk, 35, 373-374.
65. McAtee, W.L. (1921) Membracidae of the vicinity of Washington, D.C. Proceedings of the Entomological Society of Washington, 34, 123-134.
66. McKamey, S.H. (1992a) Reappraisal of the Neotropical treehopper genus Eunusa Fonseca (Homoptera: Membracidae), with ecological notes. Annals of the Entomological Society of America, 85, 253-257.
67. McKamey, S.H. (1992b) Taxonomic index to recent bibliographies of the Membracoidea. Tymbal, Auchenorrhyncha Newsletter, 19, 7-13. [includes subject index].
68. McKamey, S.H. & Deitz, L.L. (1991a) Synonymy in the treehopper genera Hoplophorion, Ochropepla, and Metcalfiella (Homoptera: Membracidae). Proceedings of the Entomological Society of Washington, 93, 159-162. [see corrections: 93, 793].
69. McKamey, S.H. & Deitz, L.L. (1991b) Nomenclatural changes in the treehopper tribes Hoplophorionini, Smiliini, and Talipedini (Homoptera: Membracidae). Proceedings of the Entomological Society of Washington, 93, 193-196. [see corrections: 93, 793].
70. McKamey, S.H. & Deitz, L.L. (1991c) Revision of the Neotropical treehopper genus Metcalfiella (Homoptera: Membracidae). North Carolina Agricultural Research Service Technical Bulletin, 294, 1-89.
71. Maddison, W.P., Donoghue, M.J. & Maddison, D.R. (1984) Outgroup analysis and parsimony. Systematic Zoology, 33, 83-103.
72. Maddison, W.P. & Maddison, D.R. (1992) MacClade, Version 3. Program and documentation. Sinauer Associates, Sunderland, Massachusetts.
73. Maddison, W.P. & Slatkin, M. (1991) Null models for the number of evolutionary steps in a character in a phylogenetic tree. Evolution, 45, 1184-1197.
74. Martorell, L.F. (1939) Insects observed in the state of Aragua, Venezuela, South America. Journal of Agriculture of the University of Puerto Rico, 23, 177-232.
75. Martorell, L.F. & García-Tudurí, J.C. (1973) Notes on the accidental introduction of Umbonia crassicornis (Amyot & Serville) - (Hemiptera: Membracidae) into Puerto Rico and its control. Journal of Agriculture of the University of Puerto Rico, 57, 307-313.
76. Mason, C.E. & Loye, J.E. (1981) An annotated list of treehoppers (Homoptera: Membracidae) of Delaware. Entomological News, 92, 33-37.
77. Masters, K.L. (1989) The adaptive significance of female-biased sex ratios in the Neotropical treehopper Umbonia ataliba (Homoptera: Membracidae). Unpublished MS thesis, University of Florida.
78. Maxwell, L.S. (1965) Florida Insects. Their Habits and Control. L.S. Maxwell, Tampa, Florida.
79. Mead, F.W. (1962) The thorn bug, Umbonia crassicornis (Amyot and Serville) (Homoptera: Membracidae). Florida Department of Agriculture. Division of Plant Industry Entomology Circular, 8, [1-2].
80. Mead, F.W. (1967) The oak treehopper, Platycotis vittata (Fabricius) (Homoptera: Membracidae). Florida Department of Agriculture. Division of Plant Industry Entomology Circular, 57, [1-2].
81. Metcalf, Z.P (1927) Homopterological gleanings no. 2. The types of certain genera of Membracidae. Entomological News, 38, 14-17.
82. Metcalf, Z.P. & Wade, V. (1963) A Bibliography of the Membracoidea and Fossil Homoptera (Homoptera: Auchenorhyncha). North Carolina State [University], Raleigh.
83. Metcalf, Z.P. & Wade, V. (1965) General Catalogue of the Homoptera. A Suppl. to Fascicle I - Membracidae of the General Catalogue of the Hemiptera. Membracoidea. In two sections. North Carolina State University, Raleigh.
84. Müller, H. (1873) Larvae of Membracis serving as milk cattle to a Brazilian species of honey-bee. Nature (London), 8, 201-202.
85. Nault, L.R., Wood, T.K. & Goff, A.M. (1974) Treehopper (Membracidae) alarm pheromones. Nature (London), 249, 387-388.
86. Nixon, K.C. & Carpenter, J.M. (1993) On outgroups. Cladistics, 9, 413-426.
87. Olmstead, K.L. & Wood, T.K. (1990) Altitudinal patterns in species richness of Neotropical treehoppers (Homoptera: Membracidae): the role of ants. Proceedings of the Entomological Society of Washington, 92, 552-560.
88. Plummer, C.C. (1936) New Membracidae (Homoptera) from Mexico, with notes on other species. Annals of the Entomological Society of America, 29, 682-693.
89. Quisenberry, S.S., Yonke, T.R. & Kopp, D.D. (1978) Key to the genera of certain immature treehoppers of Missouri with notes on their host plants (Homoptera: Membracidae). Journal of the Kansas Entomological Society, 51, 109-122.
90. Rafinesque, C.S. (1815) Analyse de la Nature ou Tableau de l'Univers et des Corps Organisés. Barravecchia, Palermo, Italy.
91. Ramos, J.A. (1979) Membracidae de la República Dominicana (Homoptera: Auchenorhyncha). University of Puerto Rico (Mayagüez), Estación Experimental Agrícola Boletín, 260, 1-71.
92. de Remes-Lenicov, A.M.M. (1978) Contribution al estudio de los membrácidos neotropicales III (Insecta-Homoptera). Neotrópica, 24, 111-116.
93. Restrepo-Mejía, R. (1980) Membrácidos de Colombia-I. Revisión parcial de las especies del género Alchisme Kirkaldy (Homoptera: Membracidae: Hoplophorioninae). Caldasia (Bogotá), 13, 103-164.
94. Richter, L. (1942) Catálogo de los membrácidos de Colombia. Revista de la Academia Colombiana, 4, 405-409.
95. Sakakibara, A.M. (1979) Micropepla mourei gen. n., sp. n. de Hoplophorioninae (Homoptera: Membracidae). Dusenia (Curitiba, Brazil), 11, 143-146.
96. Salt, G. (1929) A contribution to the ethology of the Meliponinae. Transactions of the Entomological Society of London, 77, 431-470, plates xx-xxiv.
97. Scaramuzza, L.C. (1951) Insectos observados en Eucalyptus spp. Boletín de Historia Natural de la Sociedad 'Felipe Poey', 2 (6), 86-89.
98. Schwarz, H.F. (1948) Stingless bees (Meliponidae) of the Western Hemisphere. Lestrimelitta and the following subgenera of Trigona: Trigona, Paratrigona, Schwarziana, Parapartamona, Cephalotrigona, Oxytrigona, Scaura, and Monurella. Bulletin of the American Museum of Natural History, 90, i-xviii, 1-546.
99. Stål, C. (1866) Analecta hemipterologica [continued]. Berliner Entomologische Zeitschrift, 10, 381-394.
100. Stål, C. (1869a) Hemiptera Fabriciana. Fabricianska Hemipterarter, efter de i Kopehnhamn och Kiel förvarade typexemplaren granskade och beskrifne. 2. Konliga Svenska Vetenskaps-Akademiens Handlingar, 8(1), 1-130.
101. Stål, C. (1869b) Bidrag till Membracidernas kännedom. Öfversigt af Svenska Vetenskaps - Akademiens Förhandlingar, 1869, 231-300.
102. Strümpel, H. (1972) Beitrag zur Phylogenie der Membracidae Rafinesque. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 99, 313-407.
103. Strümpel, H. (1986) Bemerkungen zur protektiven Mimese von Larven der neotropischen Membracide Leioscyta spiralis Haviland, 1925 (Homoptera: Membracidae). Entomologische Mitteilungen aus dem Zoologischen Museum, Hamburg, 8, 281-284.
104. Swofford, D.L. (1993) PAUP: Phylogenetic Analysis Using Parsimony, Version 3.1.1. Computer program distributed by the Illinois Natural History Survey, Champaign.
105. Trueman, J.W.H. (1993) Randomization confounded: a response to Carpenter. Cladistics, 9, 101-109.
106. Van Duzee, E.P. (1914) A preliminary list of the Hemiptera of San Diego County, California. Transactions of the San Diego Society of Natural History, 2, 1-57.
107. Watrous, L.E. & Wheeler, Q.D. (1981) The out-group comparison method of character analysis. Systematic Zoology, 30, 1-11.
108. Wood., T.K. (1974) Aggregating behavior of Umbonia crassicornis (Homoptera: Membracidae). Canadian Entomologist, 106, 169-173.
109. Wood., T.K. (1975) Defense in two pre-social membracids (Homoptera: Membracidae). Canadian Entomologist, 107, 1227-1231.
110. Wood., T.K. (1976a) Alarm behavior of brooding female Umbonia crassicornis (Homoptera: Membracidae). Annals of the Entomological Society of America, 69, 340-344.
111. Wood., T.K. (1976b) Biology and presocial behavior of Platycotis vittata (Homoptera: Membracidae). Annals of the Entomological Society of America, 69, 807-811.
112. Wood., T.K. (1977) Defense in Umbonia crassicornis: role of pronotum and adult aggregations (Homoptera: Membracidae). Annals of the Entomological Society of America, 70, 524-528.
113. Wood., T.K. (1979) Sociality in the Membracidae (Homoptera). Entomological Society of America. Miscellaneous Publication, 11, 15-22.
114. Wood, T.K. (1983a) Umbonia crassicornis (bicho espino, thorn bug, treehopper). Costa Rican Natural History (ed. by D.H. Janzen), pp. 773-775. University of Chicago Press, Chicago.
115. Wood, T.K. (1983b) Brooding and aggregating behavior of the treehopper, Umbonia crassicornis. National Geographic Society Research Report (Washington, D.C.), 15, 753-758.
116. Wood., T.K. (1984) Life history patterns of tropical membracids (Homoptera: Membracidae). Sociobiology, 8, 299-344.
117. Wood, T.K. (1993a) Diversity in the New World Membracidae. Annual Review of Entomology, 38, 409-435.
118. Wood, T.K. (1993b) Speciation of the Enchenopa binotata complex (Insecta: Homoptera: Membracidae). In: Evolutionary Patterns and Processes (ed. by D.R. Lees & D. Edwards), pp. 299-317. Academic Press, London.
119. Wood, T.K. & Dowell, R. (1984) Sex ratio in Umbonia crassicornis Amyot and Serville (Homoptera: Membracidae). American Midland Naturalist, 112, 58-66.
120. Wood, T.K. & Dowell, R. (1985) Reproductive behavior and dispersal in Umbonia crassicornis (Homoptera: Membracidae). Florida Entomologist, 68, 151-158.
121. Wood, T.K., Guttman, S.I. & Taylor, M.C. (1984) Mating behavior of Platycotis vittata (Fabricius) (Homoptera: Membracidae). American Midland Naturalist, 112, 305-313.
122. Wood, T.K. & Morris, G.K. (1974) Studies on the function of the membracid pronotum (Homoptera). I. Occurrence and distribution of articulated hairs. Canadian Entomologist, 106, 143-148.
123. Zak, B. (1984) Critters. Common Household and Garden Pests of Texas. Taylor Publishing, Dallas, Texas.