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note
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Supported by NSF grants EAR-9496346 (P.R.R.), EAR-9117538 (M.A.R.), and EAR-9317214 (A.R.B.), the Ann and Gordon Getty Foundation (P.R.R. and M.T.B.), and the National Natural Science Foundation of China (Z.Z.). We thank T. A. Becker and W. D. Sharp for laboratory assistance and discussions and A. L. Deino, G. H. Curtis, and an anonymous reviewer for constructive reviews of the manuscript.
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